After a meteor hits Earth, traitor forces have (for all intents and purposes) taken over, and the people are suffering. In the chaos, a boy named Tetsuro Daiba, who had lost his parents in the meteor hit, is paid a visit by Mello, a mysterious woman appearing in a bright light. When Daiba comes to, he finds itself in a spaceship infirmary under observation. Stealing a fighter that he didn`t know how to pilot (until he tried it), he breaks out, flying like a total pro. 
He was able to remember how to pilot from his DNA memory... and to find out more, his doctor friend takes him to his secret underground lab, in which he acts as a professor to find out ways to stop the traitor regime. When the forces invade to take him back, a girl named Rei Yuuki mysteriously shoots down their entire carrier with only a fighter jet! Apparently, both have mysterious powers that the enemies want, which may or may not be connected to their being the only two known survivors of the meteor impact. 
While looking for a third survivor (which turned out to be a cat), the two are taken by force to Queen Photon (ruler of the traitor regime), who claims that she is out to SAVE the Earth from those who sent the Meteor. The two don't know whether to believe her or Mello until the (telepathic) cat breaks them out of the Queen's sealed chamber, and Mello comes to straighten things out. That's when the two (er, three) of them make a choice... Stay on a planet that can't support their advanced race, or strike out into the unknown of space? 
(Source: Anime News Network)

After witnessing the death of his mother and being left for dead at the hands of Count Mecha, ten-year old Tetsuro Hoshino is awakened by a mysterious woman, Maetal. In exchange for only his company, Maetal offers him the ride of his life on the Galaxy Express 999. It is said that those who travel aboard the legendary train to the end of the line can receive a free mechanical body and with it, immortality.

Impoverished and alone, Tetsuro leaps at the opportunity and joins Maetal on a quest that will span the cosmos. As the Galaxy Express 999 makes its way across our solar system and beyond, Tetsuro is drawn deeper into a dangerous universe where peril lurks at every stop. 
(Source: Discotek)

After the death of his mother at the hands of the evil robot Count Mecha, young Tetsuro has become a delinquent street urchin who dreams only of revenge. 
Tetsuro lives in a far future world in which humans have the choice to travel to another planet where they can exchange their bodies for immortal, robotic counterparts. His goal is to someday acquire (through theft) enough money to board the Galaxy Express 999, a train which is capable of traveling through space, and which will take him to the planet of robots. 
One day, after escaping from the police, he meets the beautiful Maetel, who resembles his mother. She offers to take him on the Galaxy Express 999 to get a robotic body and to get revenge on Count Mecha. Along the journey, the two meet many varied people (including Captain Harlock and Queen Emeraldas), and Tetsuro learns what it means to be human. 
(Source: Anime News Network)

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