The second season of Sidonia no Kishi.
 
Having narrowly avoided the collision with the asteroid, the Sidonia sets a course for the Lem system in order to wipe out the Gauna nest.
 
(Source: Polygon Pictures)

It's been a thousand years since the Gauna, a strange alien race with no known method of communication, destroyed the solar system. A portion of humanity managed to escape using enormous "seed ships" like the Sidonia, which have allowed them to maintain the population while drifting through space. Nagate Tanikaze is a young man who has been raised deep in the bowels of the ship. When he goes into training to pilot the huge robotic weapons known as Gardes, Nagate is entrusted with piloting the legendary unit known as Tsugumori. Nagate and his fellow pilots put their lives on the line against the Gauna, in the ultimate battle for the survival of humanity! 
(Source: AniDB)

Outer space, the far future. 
A lone seed ship, the Sidonia, plies the void, ten centuries since the obliteration of the solar system. The massive, nearly indestructible, yet barely sentient alien life forms that destroyed humanity’s home world continue to pose an existential threat. 
Nagate Tanikaze has only known life in the vessel’s bowels deep below the sparkling strata where humans have achieved photosynthesis and new genders. Not long after he emerges from the Underground, however, the youth is bequeathed a treasured legacy by the spaceship’s coolheaded female captain. 
Meticulously drawn, peppered with clipped humor, but also unusually attentive to plot and structure for the international cult favorite, Knights of Sidonia may be Tsutomu Nihei’s most accessible work to date even as it hits notes of tragic grandeur as a hopeless struggle for survival unfolds.
 
(Source: Kodansha USA)

After the Earth was destroyed by mysterious alien lifeforms known as the Gauna, surviving remnants of the human race escaped to space in the enormous generation ship Sidonia.
Having drifted through space for millennia, the Sidonia found itself once more under attack from Gauna for the first time in a century. 
 
Once again facing the threat of extinction, a temporary victory against the Gauna was eked out thanks to the human-Gauna hybrid Tsumugi Shiraui and ace Guardian mech pilot Nagate Tanikaze. 
 
10 years later...
The people of Sidonia enjoy a brief respite.
Peaceful days pass, during which Tsumugi begins to realize her feelings for Nagate, who is now celebrated as a hero of Sidonia.
However, as Captain Kobayashi has always known, as long as the Gauna remain, peace cannot last. 
 
The decision is made: a final battle, upon which rests the fate of humanity's last survivors.
As the end approaches, will the crew be able to protect those they love? 
 
(Source: Polygon Pictures)

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