Lordgenome is the ominous Spiral King, who rules the planet from his capitol city, Teppelin. An immortal human who lived for more than 1000 years so far, he cares little for people, treating the strange girls he surrounds himself with as dolls to be cast aside when they've outlived their interest to him. From Guame we find out that Lordgenome despises human consciousness and self-awareness although we are yet to be given a reason for this dislike.  When Simon and Nia finally reach him, he claims that the act of trapping the humans underground and hunting down those who live on the surface is a means he has found to protect mankind, because according to him, when the population of humans on the surface reach one million, a terrible fate will fall upon them. He also shares the fact that he and Simon share the same "Spiral" powers, before being defeated via being stabbed in the chest by Simon's Core Drill.  In episode 18, Lordgenome's memories are retrieved by Rossiu's research team, and he reveals to Rossiu the truth about the Anti-Spirals, and the reason why he forced mankind to live underground in order to control its population. It is also revealed that at one time Lordgenome had even been part of a group that fought against the Anti-Spirals, the Spiral Knights, who fought and protected the galaxy from the Anti-Spiral menace. In the end, however, the Spiral Tribe lost the war, and the Anti-Spiral Tribe (The Han-Rasen Tribe) gained control of the universe.  Lordgenome's revived head was used to control the massive spaceship Arc-Gurren. According to Leeron, the ship only reacts to him just like the Lagann only works with Simon. After the Gurren Brigade gained control of the Cathedral Terra, it was relocated there, apparently for the same reason. Source - Wikipedia

In a far away future, mankind lives underground in huge caves, unknowing of a world above with a sky and stars. 
In the small village of Jiha, Simon, a shy boy who works as a digger discovers a strange glowing object during excavation. The enterprising Kamina, a young man with a pair of rakish sunglasses and the passion of a firey sun, befriends Simon and forms a small band of brothers, the Gurren Brigade, to escape the village and break through the ceiling of the cave to reach the surface, which few believe exist. 
The village elder won't hear of such foolishness and punishes the Brigade. However, when disaster strikes from the world above and the entire village is in jeopardy, it's up to Simon, Kamina, a girl with a big gun named Yoko, and the small yet sturdy robot, Lagann, to save the day. 
The new friends journey to the world above and find that the surface is a harsh battlefield, and it's up to them to fight back against the rampaging Beastmen to turn the tide in the humans' favor! Pierce the heavens, Gurren Lagann!

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An actor who has been in many live-action dramas.

[Synchronkartei](https://www.synchronkartei.de/sprecher/649/2)

Seven years have passed after the battle of Teppelin... Humans have since reclaimed the surface of the earth and enjoy an unprecedented period of peace and prosperity. However, humanity's increasing population growth triggers the sudden emergence of an unknown, powerful enemy. 
The fearsome, manipulative power of the mysterious Anti-Spiral proves too overwhelming for humans to even fight back. When everyone becomes desperate and loses hope, Team Dai-Gurren members reunite to stand up once again!
(Source: Official US Website)

This is the story of a man who has yet to realize what destiny holds in store for him….

In the distant future, mankind has lived quietly and restlessly underground for hundreds of years, subject to earthquakes and cave-ins.  Living in one such village are 2 young men: one named Simon who is shy and naïve, and the other named Kamina who believes in the existence of a “surface” world above their heads. The destiny of these two starts moving drastically when the ceiling of their village falls in, and a gigantic “Gunmen” and a beautiful girl named Yoko, wielding a superconductive rifle, come from the surface. Together, Kamina, Simon and Yoko ride the mecha “Lagann” that Simon digs out of the ground, and fly up to the surface!

However, the surface is not the dreamland that Kamina envisioned. This world is ruled by the Spiral King and his army of beastmen. Kamina and Simon, along with their comrades, challenge the Spiral King in an attempt to change a desperate world to one full of hope for the future with Gurren Lagann!

In their closed-off underground village, Kamina and Simon chafe at the limits imposed by the village elder. Yet all this will change, when Simon stumbles across a fantastic device - just as the village's peace is broken by a violent intrusion.


A re-imagining of Gurren Lagann. In this alternate version of TTGL, Simon lives in Tepelen city and dreams of becoming a Gunmen pilot after having seen the stars through a crack in the sky. One day, he comes across a strange small Gunmen. The hatch opens, and out comes a beautiful girl named Nia.


Tengen Toppa Gurren Lagann: Gurren Gakuen-Hen (Gurren School Edition) will feature the cast of Gainax's robot mecha anime in a school in a parallel world in multidimensional space. As the character Kamina puts it, the cast will "enjoy school life" and "sweat under the glittering sun."
 
(Source: Anime News Network)